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I hear the forerunner 205 had satelite issues. I've been using the 305 for the last year or so and haven't had any problems running in the city, or on the trails. I've even used it geo-caching and hiking as a GPS. You can't load maps on it, but you can mark plenty of waypoints.
The virtual partner is by far the best feature for exercise though. You can have a computer simulated partner based on times you want to run over-all, or you can create a virtual partner that mimics your past (or best) runs. This way you know what pace you should keep on parts of the course that may be inclines or declines based on how you performed on those parts before.
I haven't done my research on the 405 as I didn't even know it existed, but I may have to look into it.
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